This is Part I of an article that was
published in the October, 2003 issue of Orchids Australia.
Where are we coming from? Where are we headed?
Yellow phalaenopsis breeding –
an overview
Aesop tells us the story of the
race between the tortoise and the hare. The hare was very fast but only
ran in fits and starts and never completed the race while the tortoise was
much slower and steadier and kept plodding until he eventually won.
Although there is no race, the two major branches of novelty phalaenopsis
breeding can be compared to the two participants in this race.
Like the hare, red breeding has
gone in fits and starts. Until the mid-1990s, almost all red breeding
resulted in a few, generally fragrant red flowers. Flower count (per
inflorescence) was usually quite low, shape was starry and size was
relatively small. The size often remained in the 2-2½-inch range and most
new hybrids just produced more of the same. Every time a
"spectacular" red flower came along, there was a spurt of
breeding with it. Phal. Luedde-violacea, Phal. George Vasquez and Phal.
Golden Buddha have each been the cause for such spurts of growth. Each
of these plants continues to be used but to a lesser extent. However, in
the mid- and late-1990s, there was an unprecedented improvement in red
breeding (courtesy almost exclusively of Phal. Brother Purple), and
this virtually overnight. Reds achieved a degree of excellence never
before reached. While we await the next "star" in the firmament
of red breeding, Phal. Brother Purple and its progeny are enjoying
their time in the sun.
Yellow phalaenopsis breeding, on
the other hand, began in earnest about 40 years ago and, like the
tortoise, has progressed on a slow and steady pace. There have been
periods that produced progress more rapidly than others and along the way
there have been several important stud plants but the progress has
generally been slow, steady and ongoing. This article will discuss yellow
breeding, focusing on the most important stud plants.
It is hard to examine plant
breeding in a "logical" manner for various reasons. The most
important is the fact that plant breeding, unlike human breeding, is not
necessarily either chronological or linear. In plant breeding, it is
common to attempt to reinforce traits by trying to produce homozygous
genes through inbreeding or linebreeding. This may be done in many ways,
including breeding two plants together with a common parent, breeding a
plant back to a species in its background, breeding a plant back to its
parent or through sibling crosses.
The first yellow phalaenopsis
hybrid registered was Phal. Golden Louis, registered by Lewis
Vaughan in 1957. It was a hybrid of Phal. Doris and Phal.
mannii. (It was originally thought that there were three white
Phalaenopsis species: Phal. amabilis, Phal. aphrodite and
Phal. rimestadiana, used to create Phal. Doris. Some
taxonomists now consider that all three are at most 2 species, Phal. amabilis
and Phal. aphrodite and some people believe that Phal. Doris
is simply a "super" Phal. amabilis.)
More yellow hybrids with Phal.
mannii followed Phal. Golden Louis since Phal. mannii
was one of the first yellow species in common cultivation. The standard
for judging Phalaenopsis emphasizes roundness, fullness and flatness. Many
people have said semi-jokingly that Phal. mannii was a perfect
breeder’s plant: no matter what you bred it to, the result was an
improvement. This may be quite true. Although there are over 130
registered first-generation hybrids with Phal. mannii, only two
could be considered reasonably well known: Phal. Su-An Cricket (x Phal.
Gelblieber) and Phal. Mambo (x Phal. amboinensis).
Neither of these two hybrids is round nor full.
Several other species have been
used in yellow breeding. The most important have been Phal. fasciata,
Phal. amboinensis, Phal. venosa, Phal. lueddemanniana
and, to a much lesser extent, Phal. violacea and Phal. gigantea.
Phal. gigantea,
in much the same way as Phal. amboinensis, comes in two varieties:
red spotting over a cream base color or over a yellow base color. Its
influence in yellow breeding is seen almost exclusively in Phal. Liu
Tuen-Shen (see below). As for Phal. violacea, the species that was
used in breeding was generally the variety from Borneo, now called Phal.
bellina, although the RHS still calls it Phal. violacea for
registration purposes. As a rule, Phal. violacea has been used in
breeding for reds. What influence it has had in yellow breeding is
generally as a result of "failed" red breeding.
The story of Phal. fasciata
breeding is similar to that of Phal. mannii in that very few of its
progeny are well known despite the fact that well over 100
first-generation hybrids have been registered. The best known of these are
all primary hybrids: Dtps. Red Elf (x Dor. pulcherrima), Phal.
Golden Pride (x Phal. amboinensis) and Phal. Spica (x Phal.
lueddemanniana). Dtps. Red Elf was made by Hermann Pigors and
registered by Lenette Nurseries. One particular clone ‘Lenette#4’
received an HCC and has been widely distributed. H. P. Norton
started a whole new line of red breeding using this (and other) hybrids of
Dor. pulcherrima but fertility problems persist and, in the long
run, this line of breeding will probably be little more than a flash in
the pan. Phal. Golden Pride appeared to have great potential in its
day and, its most well known offspring, Phal. Solar Flare (x Phal.
Golden Sands), did manage to garner 28(!) awards but, by today’s
standards, we see that the flowers are too few and too clustered at the
ends of the inflorescences.
Phal. Spica
is the most important Phal. fasciata (and Phal. lueddemanniana)
hybrid in yellow breeding. Registered by Lee Kuhn (the founder with wife
Janet of J & L Orchids) in 1969, it has gone on to produce two of the
best known and most important stud plants in novelty breeding, Phal. Barbara
Moler and Phal. Golden Buddha. Both of these plants have the two
most important traits for hybridizers – they are (or were) readily
available and both are very fertile. Most of their offspring are quite
fertile as well. The legacy of Phal. Spica will live on in yellow
breeding through many of its most important progeny: Phal. Misty
Green (Barbara Moler x Bamboo Baby), Phal. Golden Gift (Golden
Buddha x Deventeriana), Phal. Fortune Buddha (Golden Buddha x Liu
Tuen-Shen) as well as Phal. Baldan’s Kaleidoscope (Hausermann’s
Candy x Daryl Lockhart). Because many of the Phal. Golden Buddha
hybrids concentrated on red progeny, the legacy of Phal. Spica is
even more important in the context of red breeding.
The most famous and the most
influential Phal. fasciata hybrid was not registered with Phal.
fasciata as the parent. It is Phal. Golden Sands. Roy Fields
registered it in 1964 as Phal. Fenton Davis Avant x Phal.
lueddemanniana var. ochracea. Despite the official
registration, "[t]here seems to be no question that Roy Fields used a
Phalaenopsis fasciata and not Phalaenopsis lueddemanniana
v. ochracea to breed the cross." Phal. Golden Sands is
discussed below.
Phal. lueddemanniana
is one of the most versatile of the phalaenopsis species with several very
(optically) different varieties. Depending on whether you are a lumper or
a splitter, you may consider each variety to be a separate species.
However, with the exception of Phal. Spica (see supra), most
of the well-known hybrids of Phal. lueddemanniana, with the
exception of Phal. Tyler Carlson (x Phal. Gladys Read), all
produced spotted or red flowers: Phal. Paifang’s Queen (x Phal.
Mount Kaala), Phal. Coral Isles (x Phal. Princess
Kaiulani), Phal. Cabrillo Star (x Phal. Ramona), Phal. Zadian
(x Phal. Zada) and Phal. Luedde-violacea (x Phal. violacea).
The reason for this is quite simple. As a rule, because of its color, Phal.
lueddemanniana var. pulchra was the commonly used variety in breeding.
When bred to a red flower, more red flowers were produced and, when bred
to a flower with less coloring, the "solid" red of the Phal.
lueddemanniana var. pulchra broke down into its component spots and
bars.
Phal. amboinensis
is possibly the most important species in
yellow breeding to date. However, the Phal. amboinensis that was
originally discovered was cream with brown barring. It was used in
hybridizing but it wasn’t until the 1950s that the yellow form of the
species was discovered and in the ’60s that it was widely cultivated.
Then, Phal. amboinensis really came into its own. It was bred to
almost anything and produced hundreds of well-known and well-used progeny.
A list of awarded progeny could take pages but some of the more important
would definitely include Phal. Sarah Frances Pridgen (x Phal. Barbara
Moler), Phal. Summer Wine (x Phal. Bettylee Burke), Phal.
Ambo Buddha (x Phal. Brother Buddha), Phal. Paula
Hausermann (x Phal. Kenneth Stromsland), Phal. Sara Lee (x Phal.
Princess Lorraine), Phal. Brother Horace (x Phal. Brother
Glamour), Phal. Penang (x Phal. Rosy Charm), Phal. Malibu
Imp (x Phal. Luedde-violacea) and Phal. Brother Elizabeth (x
Phal. Brother Cucumber).
With the one notable exception of
Phal. Bamboo Baby, almost all of the best-known hybrids of Phal.
amboinensis are its primary hybrids: Phal. Corona (x Phal.
cornu-cervi), Phal. Deventeriana (x Phal. amabilis), Phal.
Golden Pride (x Phal. fasciata), Phal. David Lim (x Phal.
gigantea), Phal. Ambomanniana (x Phal. lueddemanniana), Phal.
Mambo (x Phal. mannii), Phal. Ambotrana (x Phal.
sumatrana), Phal. Ambonosa (x Phal. venosa), Phal. Ambotris
(x Phal. equestris) and Phal. Princess Kaiulani (x Phal.
violacea). However, despite their notoriety, very few of these primary
hybrids, with the exception of Phal. Deventeriana and Phal. Princess
Kaiulani, have to date produced anything of long-term note.
Phal. venosa
(originally called Phal. psilantha and awarded as such) was only
discovered about 20 years ago. It sparked a great deal of interest as many
people considered it to be a worthy successor and possibly an even better
and more important species for breeding yellows than Phal. amboinensis.
The first hybrids made with Phal. venosa were only registered in
1984, a little over 15 years ago. Several hybridizers remade hybrids that
had been made with Phal. amboinensis. Others struck out in new and
different directions. However, many people now freely admit that breeding
with Phal. venosa can be difficult and that, while Phal. venosa
hybrids can be very attractive, breeding with some of them is nearly
impossible. For example, Phal. Zuma Aussie Delight (Sweet Memory x venosa)
has been meristemmed and is in hundreds, if not thousands, of collections.
Despite this fact, there are only a couple of registered hybrids and
thousands of failed attempts.
Phal. venosa
was used in making several primary hybrids including Phal. Kuntrarti
Rarashati (x Phal. equestris). To date, Phal. Kuntrarti
Rarashati has produced some of the nicest and most interesting multiflora
hybrids around – Phal. Taida Pixie (x Phal. Carmela’s
Pixie), Dtps. Pico Lady Ruby (x Dor. pulcherrima) with one
Award of Quality and several flower quality awards, Phal. Jori
Moore (x Phal. Zuma’s Pixie) which also has both an Award of
Quality and awarded progeny as well as Phal. Bedford Baby (x Phal.
amboinensis), with one flower quality award to date. Phal. Penang
Girl (x Phal. violacea), registered in 1984 has been used
approximately 30 times to date in making registered hybrids. It is both
colorful and fertile, a wonderful combination. A delightful fragrance is
an added bonus. Phal. Ambonosa (x Phal. amboinensis) has
been used extensively in breeding. There are several awarded clones with
flowers ranging from about 4.5 to 5.5 cm natural spread and the flowers
can be quite plentiful because of the tendency to rebloom on old
inflorescences. It is the parent of about 30 registered hybrids to date,
having been used since 1989. Most of the hybrids of Phal. Ambonosa
to date are hardly known but Phal. Minho Queen (x Phal. Paifang’s
Auckland) has one award and Phal. Brother Torro (x Phal. Peach
Boy) has several.
There is a group of Phal.
venosa hybrids that can best be lumped as near-primary hybrids. There
are many of these including Phal. John Ewing (x Phal. George
Vasquez) and Phal. Venimp (x Phal. Malibu Imp) in the United
States and Phal. Coral Nosa (x Phal. Coral Isles) in Taiwan.
The flowers in this group all tend to be quite dark, star-shaped and few
in number. However, they are all fragrant and relatively new to the
hybridizing world. Of these, Phal. Coral Nosa has probably been the
plant that has been used the most in hybridizing. It is the parent of Phal.
Black Eagle (x Phal. George Vasquez), Phal. Chiayi
Golden Moon (x Phal. Maritea), Phal. Ta Lin Kaiulani (x Phal.
Princess Kaiulani) and Phal. Black Beauty (x Phal. gigantea),
all of which have played important roles in phalaenopsis breeding in
Taiwan.
Phal. Golden
Bells (x Phal. Golden Sands), a hybrid registered in 1988 by H. P.
Norton, is one of the most prolific Phal. venosa hybrids. It has
received three awards to date and is the parent of almost 90 registered
hybrids to date. In spite of the relative youth of this grex, it has
already begun to prove its worth with several of its first-generation
hybrids such as Phal. Orchidview Bellringer (x Phal. Deventeriana),
Phal. Melodious Bells (x Phal. Golden Buddha) and Phal. Golden
Circles (x Phal. Misty Green) but, yet again, it is much too early
to examine the complete impact of this grex.
Some of the most important
(commercially, anyway) hybrids of Phal. venosa are crosses with
large whites. This type of breeding was done with Phal. amboinensis
and the tradition continues with Phal. venosa. As a rule, this
produces good flower count with yellow flowers that are reasonably large.